FLORA OF THE VEDAUWOO NATURAL HISTORY SITE (VNHS)
The following alphabetic list, by family, reflects species currently
in the LCCC herbarium or listed on the College (Satellite) Mountain
Land Inventory that was completed by LCCC faculty in 1984 (unpublished).
Family decriptions are taken from Porter (1968) and Dorn (1988).
Page number references following each family name correspond to
the text "Vascular Plants of Wyoming" (Dorn 1988). It is strongly
recommended that one consult this source when attempting to identify
plants to the species level.
Go back to: VNHS General Info., Botany & Ecology Homepage at NWU, Department of Biology Homepage, Nebraska Wesleyan University Homepage.
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QUICK REFERENCE FOR PLANT FAMILIES AT VNHS
1. ACERACEAE Maple Family (page 30)
Trees or shrubs; opposite leaves, either simple and palmately
lobed or compound and pinnate; flowers small, regular; sepals
5; petals usually 5 or none; stamens 3-12; pistil 1, of 2 weakly
united carpels; ovary superior; fruit a paired samara, the wings
with prominent veins.
Acer glabrum Torr. - Rocky Mtn. Maple
2. APIACEAE Parsnip Family (page 32)
Aromatic herbs; leaves alternate or basal, usually compound having
petioles dilated and sheathing the stem; flowers small, usually
in compound umbels, epigynous, without a hypanthium, regular or
somewhat irregular; sepals 5, sometimes minute or obsolete; petals
5, usually white or yellow; stamens 5; pistil 1, of 2 united carpels;
styles 2; ovary inferior, 2-celled and 2-seeded; fruit a schizocarp
(the 2 carpels separating at maturity into 2 1-seeded mericarps).
Angelica ampla - Ample Angelica
Conioselinum scopulorum - Rocky Mountain Hemlock Parsley
Cymopterus lemmonii - Lemmon Spring Parsley
Harbouria trachypleura (Gray) Coult. &Rose - Harbouria
Heracleum sphondylium (L.) Michx. Dorn var. lanataum - Hogweed Cowparsnip
Lomatium orientale Coult. &Rose - Biscuitroot
Osmorhiza chilensis - Chilean Sweet Cicely
Osmorhiza depauperata Phil. - Long Style Sweetroot
Sanicula marilandica L. - Black Sanicle
3. AMARANTHACEAE Pigweed Family (page 31)
Annuals; leaves simple, usually entire, alternate or opposite;
flowers bisexual or unisexual, regular or slightly irregular,
usually subtended by scarious bracts, in racemes, spikes, or axillary
clusters; sepals 3-5, 1-4 mm long, usually dry and membranous,
separate or not; petals none; stamens 1-5, the filaments often
united into a tube; ovary 1, superior; styles 1-3; fruit usually
a utricle or nutlet.
Amaranthus blitoides - Creeping Pigweed
4. APOCYNACEAE Dogbane Family (page 41)
Rhizomatous herbs, juice milky; leaves simple, opposite, entire;
flowers bisexual, regular, in cymes; sepals 5, united; petals
5, united; stamens 5, alternate with corolla lobes and attached
to tube; ovaries 2, superior or nearly so; style 1, very short
and enlarged; fruit a follicle.
Apocynum medium Green - Dogbane
5. ASTERACEAE Sunflower Family (page 43)
Herbs or shrubs of various aspects; flowers in dense heads on
a common receptacle that is surrounded below by an involucre of
several to many bracts, each head often appearing like a single
flower, epigynous, quite small, with united petals, all tubular,
all ligulate (with strap-shaped corollas that are usually conspicuous
and larger than the tubular ones), or the central (disk) flowers
with tubular corollas and the marginal (ray) flowers with ligulate
corollas; stamens 5, attached to the corolla tube and usually
united by their anthers into a ring through which the style is
projected; pistil 1; ovary inferior; style single but cleft into
2 stigmas at the summit; fruit from each flower (if fertile) an
achene that is crowned by a late-developing calyx (the pappus)
composed of hairs, bristles, awns, or scales.
Achillea millefolium L. - Common Yarrow
Agoseris glauca var. lacinata (Eaton) Smiley - False Dandelion
Antennaria anaphaloides Rybd. - Pearly Pussytoes
Antennaria microphylla Rydb. - Littleleaf Pussytoes
Antennaria parvifolia Nutt. - Smallleaf Pussytoes
Arnica cordifolia Hook. - Heartleaf Arnica
Arnica fulgens - Gleaming Arnica
Artemisia campestris - Northern Sagewort
Artemisia dracunculus - Tarragon Sagewort
Artemisia frigida Willd. - Fringed Sagewort
Artemisia ludoviciana - Louisiana Sagewort
Artemisia tridentata - Big Sagebrush
Artemisia tripartita - Three-lip Sagebrush
Aster glucodes - Glaucous Aster
Aster laevis - Smooth Aster
Bahia dissecta (Gray) Britt. - Ragleag Bahia
Brickellia grandiflora (Hook.) Nutt. - Tasselflower Brickellbush
Chrysothamnus viscidiflorus - Douglas Rabbitbrush
Cirsium arvense - Thistle
Cirsium canescens Nutt. - Thistle
Cirsium pulcherrimum - Beautiful Thistle
Crepis acuminata Nutt. - Tapertip Hawksbeard
Crepis modocensis - Yellowstone Hawksbeard
Erigeron caespitosus - Caespitose Fleabane
Erigeron compositus Pursh var. discoideus Gray - Fernleaf Fleabane
Erigeron flagellaris Gray - Trailing Fleabane
Erigeron pumulis Nutt. - Low Fleabane
Erigeron speciousus - Showy Fleabane
Erigeron vetensis - Glandular Fleabane
Gaillardia aristata Pursh - Common Gaillardia
Grindelia squarrosa - Common Gumweed
Grindelia subalpina Greene - Gumweed
Helianthus petiolaris - Prairie Sunflower
Helianthus pumilus Nutt. - Sunflower
Heterotheca villosa (Pursh) Shinners var. villosa - Telegraph plant
Hymenoxys acaulis (Pursh) Parker - Stemless Actinea
Liatris punctata Hook. - Dotted Grayfeather
Machaeranthera linearis Greene
Rudbeckia laciniata - Laciniate Coneflower
Senecio canus - Wooly Groundsel
Senecio fendleri Gray - Fendler Groundsel
Senecio hydrophilus - Water Groundsel
Senecio integerrimus Nutt. var. exaltatus (Nutt.) Cronq. - Lambstounge Groundel
Senecio rapifolius - Rock Groundsel
Senecio spartioides var. fremontii (T.?.);Greene ex Ulms - Broom Groundsel
Solidago canadensis - Canada Goldenrod
Solidago missouriensis Nutt. var. fasculuata Holz. - Missouri Goldenrod
Taraxacum officinale Weber. - Dandelion
Townsendia hookeri Beaman - Hoary Townsendia
Tragopogon dubius Scop. - Yellow Salsify
6. BERBERIDACEAE Barberry Family (page 87)
Low shrubs; leaves alternate, pinnately compound, leathery; leaflets
spine-toothed; flowers small, yellow, hypogynous, regular; sepals
6; petals 6; stamens 6, opposite the petals, the anthers opening
by terminal valves; pistil 1, becoming fruit a 1-3 seeded berry
with a bluish color fruit.
Mahonia repens (Lindl.) G. Don - Barberry
7. BETULACEAE Birch Family (page 88)
Trees or shrubs; leaves alternate, simple, often doubly serrated
and sometimes lobed; flowers unisexual, at least the male flowers
produced in catkins and sometimes that female flowers also in
catkins, both male and female catkins produced on the same plant,
the plants thus monoecious; fruit a small nutlet, with or without
wings, or sometimes (inCorylus, the Hazelnut of northeastern Wyoming) a larger nut enclosed
in united, leafy bracts of an involucre.
Alnus incana L. - Alder
8. BORAGINACEAE Borage Family (page 89)
Smooth or bristly-hairy herbs; leaves alternate, entire; flowers
regular, hypogynous; sepals 5, united below; petals 5, united
below into a slender or funnelform tube, often with appendages
at the throat of the corolla forming a corona; stamens 5, attached
to the tube of the corolla alternate with its lobes; pistil 1;
ovary deeply 4-lobed; style 1, arising from between the ovary
lobes; fruit a group of 1-seeded nutlets, or sometimes a single,
bone-like nutlet.
Cryptantha celosioides - Northern Miner's Candle
Cryptantha virgata (Porter) Pays. - Cryptantha
Cynoglossum officianle L. - Common Houndstounge
Hackelia floribunda (Lehm.) Johnst. - Western Tickweed
Lappula redowskii (Hernam.) Greene var. redowskii - Blueburr Stickseed
Mertensia ciliata (James ex Torr.) G. Don - Mountain Bluebells
Mertensia lanceolata (Pursh) D.C. ex A. DC - Lanceleaf Bluebells
9. BRASSICACEAE Mustard Family (page 95)
Annual or perennial, erect or caespitose herbs with watery and
often peppery juice; flowers hypogynous, 4-merous, regular, produced
in racemes; sepals 4; petals 4, often with long claws; stamens
usually 6, the outer 2 stamens usually 6, the outer 2 stamens
short and the inner 4 stamens longer but included in the corolla,
and sometimes the stamens only 2; pistil 1, of 2 united carpels
joined by a delicate partition that remains on the plant when
the fruit split apart; fruit a silique (a rather long fruit) or
a silicle (a globose or very short silique); seeds 1 or more in
each carpel.
Alyssum desertorum Stapf. - Pale Alyssum
Arabis drummondii Gray - Drummond Rockcress
Arabis glabra - Glabrous Rock Cress
Arabis microphylla - Little-Leaved Rock Cress
Camelina microcarpa Andrz. ex D.C. - Littleseed Falseflax
Descurainia pinnata - Pinnate Tansy Mustard
Descurainia sophia (L.) Webb ex Prantl. - Flixweed Tansymustard
Draba nemorosa L. - Woods Draba
Draba streptocarpa Gray - Draba
Erysimum inconspicuum - Inconspicuous Wallflower
Erysumum asperum var. arkansum (Nutt.) Gray - Plains Wallflower
Lepidium densiflorum Schrad. - Dense-Flowered Pepper Grass
Lepidium ramosissimum A. Nels - Branched Pepperweed
Lesquerella montana (Gray) Wats. - Mountain Bladderpod
Sisymbrium linifolium - Linear-Leaved Hedge Mustard
Thlaspi arvense L. - Penneycress
Thlaspi montanum L. - Alps Pennycress
10. CACTACEAE Cactus Family (page 115)
Fleshy stemmed, branched, or unbranched, usually spiny perennials,
the leaves in ours none or small, fleshy, linear, and produced
only on juvenile shoots and early deciduous; branches, flowers,
and spines produced from cushionlike structures called areoles;
stems flat or rounded and jointed, or subglobose to cylindric
and with ribs or tubercles or both; flowers with several sets
of perianth parts, yellow, green, pink, or purple, epigynous,
perfect, and regular; stamens numerous; pistil 1, of several united
carpels; ovary 1-celled, parietal placentation; style 1; stigmas
several; fruit fleshy or dry, with several to many seeds, splitting
open regularly or indehiscent.
Opuntia polycantha - Plains Pricklypear
Pediocactus simpsonii - Simpson Barrel Cactus
11. CALLITRICHACEAE Water Starwort Family (page 116)
Aquatic herbs; leaves simple, opposite, entire; flowers axillary,
the 2 sexes often close together to appear like a single flower,
monoecious; perianth none (2 bracts present); stamen 1; ovary
1; styles 2; fruit a schizocarp, splitting into 2 or 4 segments
with 1 seed each.
Callitriche palustris - Palustrine Water Starwort
12. CAMPANULACEAE Bellflower Family (page 117)
Herbs with milky juice; leaves alternate; flowers epigynous; sepals
5; petals 5, united to form a campanulate (bell-shaped) corolla,
regular, blue; stamens 5, free form the corolla; filaments broadened
at the base; pistil 1; style single; stigmas 3; ovary inferior,
3-celled; fruit a capsule.
Campanula rotundifolia L. - Bluebell Bellflower
13. CANNABACEAE Hemp Family (page 118)
Dieocious vines; leaves alternate or opposite, simple; flowers
mostly regular staminate in racemes or panicles, pistillate in
dense clusters; sepals 5, all joined to form a cup in pistillate
flowers; petals none; stamens 5; pistil 1; ovary superior; styles
1 or 2; fruit an achene.
Humulus lupulus L. - Woodbind, Wild Hops
14. CAPRIFOLIACEAE Honeysuckle Family (page 119)
Shrubs; leaves opposite, simple or compound; flowers epigynous,
regular to somewhat irregular; sepals 4-5; petals 4-5, united
into a short or long tube below; stamens as many as the lobes
of the corolla, attached to the corolla tube alternate with its
lobes; pistil 1; ovary inferior, 2-5 celled; fruit a berry, sometimes
drupaceous or a capsule.
Lonicera involucrata (Richards.) Banks ex Spreng. - Bearberry Honeysuckle
Symphoricarpus albus L. - Common Snowberry
15. CARYOPHYLLACEAE Pink Family (page 120)
Herbs, not succulent; leaves opposite, entire; flowers usually
in cymes; sepals 4-5, distinct or united below; petals none or
usually 4-5; stamens the same number as the sepals or twice as
many; pistil 1, of 2-5 united carpel; ovary superior, 1-celled;
placentation free-central; styles 2-5; fruit a capsule with a
few to many seeds.
Arenaria fendleri Gray - Fendler Sandwort
Cerastium arvense L. - Mouse-eared Chickweed
Lychnis drummondii - Drummond Campion
Paronychia depressa (T. &G.) Nutt. ex A. Nels. - Nailwort
Paronychia sessiliflora - Sessil-Flowered Whitlow-Wort
Silene noctiflora - Night-Flower Catchfly
Stellaria longifolia Muhl. ex Willd. - Longstock Starwort
16. CHENOPODIACEAE Goosefoot Family (page 127)
Herbs or shrubs; leaves simple, alternate, rarely opposite, sometimes
succulent or gray-scurfy; flowers bisexual, or unisexual and the
plants monoecious or dioecious, regular or rarely irregular, solitary
or clustered in leaf axils or in spikes, racemes, or panicles;
sepals usually 5, rarely fewer, usually united, sometimes none;
petals none; stamens usually 5, rarely fewer; ovary 1, superior;
styles or stigmas 2 or 3; fruit an achene or utricle.
Chenopodium berlanderi - Berlandier Goosefoot
Chenopodium fremontii - Fremont Goosefoot
Chenopodium glaucum - Glaucous Goosefoot
Monolepis nuttalliana (Schult.) Greene - Nuttall Monolepis
17. COMMELINACEAE Spiderwort Family (page 134)
Fibrous rooted herbs; leaves simple, lance-linear or linear, alternate,
sheathing; flowers bisexual, mostly regular, in a cyme or umbel
or solitary; sepals 3, green, separate; petals 3, blue, usually
separate; stamens 6, filaments usually villous; pistil 1; ovary
superior; style 1; locules 1-3; ovules 1 or few per locule; fruit
a capsule.
Tradescantia occidentalis - Western Spiderwort
18. CORNACEAE Dogwood Family (page 135)
Shrubs or perennial herbs; leaves simple, entire, opposite or
apparently whorled; flowers usually bisexual, regular, in cymes
or a flower-like, whitish, involucrate head; sepals 4, united;
petals 4, separate, white, greenish-white, or purplish; stamens
4; ovary 1, inferior; style 1; locules 2; ovules 1 per locule;
fruit a red, white, or bluish drupe.
Cornus stolonifera Michx. - Redosier Dogwood
19. CRASSULACEAE Stonecrop Family (page 135)
Fleshy, succulent herbs; leaves simple, entire, sessile; flowers
hypogynous, yellow, pink, or purplish; sepals 4-5; petals 4-5;
stamens twice as many as the sepals or petals; pistil (carpels)
4-5; fruit a group of follicles.
Sedum lanceolata Torrey - Stonecrop
20. CUPRESSACEAE Cedar Family (page 27)
Evergreen trees or shrubs; leaves scale-like or awl-shaped; cones
fleshy or leathery and berry-like cones; seed wingless, one or
more in each cone.
Juniperus communis - Common Juniper
21. CYPERACEAE Sedge Family (page 136)
Grass-like or rush-like herbs; leaves simple, alternate or basal,
sheathing, usually 3 ranked, rarely reduced to sheath; ligule
often lacking; flowers bisexual or unisexual, arranged in spikelets
or spikes, each flower subtended by a chaffy bract; perianth bristly
or scaly or absent; stamens 1-3, rarely 6; ovary 1, superior;
style 1, usually branched above; fruit an achene, enclosed in
a sac (perigynium) inCarex.
Carex aquatilis Wahl. - Water Sedge
Carex aurea - Golden Sedge
Carex filifolia Nutt. - Threadleaf Sedge
Carex geyeri - Elk Sedge
Carex lanuginosa - Woolly Sedge
Carex microptera - Little-Wing Sedge
Carex nebrascensis Dewey - Nebraska Sedge
Carex oreocharis Holm. - Sedge
Carex pachystachya - Thickspike Sedge
Carex pensylvanica - Pennsylvania Sedge
Carex stenophylla Wahl. - Sedge
Eleocharis palustris - Water Spike Rush
22. EQUISETACEAE Horsetail Family (page 22)
Rush-like herbs; stems branched or unbranched, jointed, longitudinally
striate, hollow between the joints; leaves consisting of a ring
of teeth at each joint; reproduction by spores that are all alike
and produced in a terminal cone made up of whorls of sporophylls.
Equisetum arvense L. - Horsetail
Equisetum hyemale L. - Scouring Rush, Horsetail
23. ERICACEAE Heath Family (page 151)
Herbs, shrubs, or trees; leaves mostly leathery, often evergreen,
simple, entire or obscurely toothed; flowers hypogynous or epigynous;
sepals 4 or 5; petals 4 or 5, usually united but sometimes only
slightly so, the corolla often urn-shaped; stamens as many as
the corolla lobes or twice as many, free from the corolla; anthers
often opening by terminal pores; pistil 1; carpels 2-5, united;
fruit a capsule or a berry.
Arctostaphylos uva-ursi (L.) Spreng. - Bearberry, Manzanita
Pterospora andromedea Nutt. - Woodland Pinedrops
24. EUPHORBIACEAE Spurge Family (page 154)
Flowers in perianth-like involucres; 1 stalked pistil; stamens
several to many in each involucre; juice usually milky.
Euphorbia brachycera Engelm. var. robusta Dorn - Robust Spurge
25. FABACEAE Pea Family (page 156)
Trees, shrubs, or mostly herbs; leaves alternate, usually compound,
with prominent stipules; flowers irregular, more or less perigynous
but the hypanthium short or shallow; sepals 5, united below to
form a tube; petals 5, dissimilar in size and shape, the upper
petal (standard or banner) the largest, the two lateral petals
(wings) oblique, and the two lower petals (keel) united by their
lower edges and enclosing that stamens and pistil; stamens usually
10, often diadelphous (9 united by their filaments and 1 separate),
sometimes monadelphous (all united by their filaments), and sometimes
all separate (distinct); anthers usually alike but sometimes 5
anthers larger that the other 5; pistil 1, of 1 carpel; fruit
a legume, usually leathery but sometimes papery and inflated and
sometimes fleshy.
Astragalus adsurgens Pall. var. robustior Hook. - Standing Milkvetch
Astragalus agrestis Dougl. ex. G. Don. - Field Milkvetch
Astragalus crassicarpus Nutt. - Groundlpum Milkvetch
Astragalus flesuosus (Hook.) Dougl. ex. G. Don. - Pliant Milkvetch
Astragalus shortianus Nutt. - Shorts Milkvetch
Astragulus eucosmus - Elegant Milkvetch
Astragulus parryi - Parry Milkvetch
Astragulus purshii - Pursh Milkvetch
Lupinus argenteus Pursh - Silvery Lupine
Melitolus alba - White Sweet Clover
Melitolus officinalis - Yellow Sweet Clover
Oxytropis lambertii Pursh - Lambert Loco
Oxytropis multiceps Nutt. - Flowery Loco
Oxytropis sericea Nutt. var. spicata (Hook.) Barneby - Silky Loco
26. FUMARIACEAE Fumitory Family (page 177)
Delicate, smooth herbs; juice watery; leaves compound and dissected,
fern-like; flowers yellow, hypogynous, irregular, produced in
racemes among the leaves; sepals 2, small and scale-like; petals
4, in 2 pairs, the outer pair with spreading tips and the one
or both of them spurred or saccate at the base, the inner pair
smaller; stamens 6, in 2 sets of 3 each, placed opposite the larger
petals; pistil 1, becoming a 1-celled, several-seeded, elongate
capsule in fruit; seeds black and shiny.
Corydalis aurea Willd. var. aurea - Golden Corydalis
27. GERANIACEAE Geranium Family (page 179)
Herbs; leaves palmately lobed or pinnately compound; flowers small
or large and showy, white or pink, to purple; sepals 5; petals
5, dropping early; stamens the same numbers as the sepals or petals,
or twice as many; filaments somewhat united at the base; pistil
1, of 5 united carpels that separate elastically; pistil and immature
fruit long-beaked.
Geranium caespitosum James - Pineywoods Geranium
Geranium fremontii - Fremont Geranium
Geranium richardsonii Fisch &Trautv. - Richardson Geranium
Geranium viscosissimum - Sticky Geranium
28. GROSSULARIACEAE Currant Family (page 180)
Shrubs; leaves alternate, palmately veined, palmately lobed; flowers
epigynous, with a hypanthium originating from the top of the ovary;
sepals 5, often petaloid and conspicuous; petals 5, small and
inconspicuous; stamens 5; fruit a berry with few seeds.
Ribes cereum Dougl. var. pediceilare - Wax Current
Ribes inerme - Spiny Currant
29. HYDRANGEACEAE Hydrangea Family (page 181)
Shrubs; leaves opposite, simple, without stipules; flowers white,
on the ends of the branches, perigynous, the sepals, petals, and
stamens inserted on the rim of a hypanthium; Sepals 5; petals
5; stamens 10 or sometimes more; pistil of 2-10 united carpels,
each with several seeds; fruit a capsule.
Jamesia americana T. &G. - Cliff Jamesia
30. HYDROPHYLLACEAE Waterleaf Family (page 182)
Annual or perennial herbs; leaves alternate; flowers often coiled
inflorescences, usually white or blue to purple, regular, hypogynous,
with the united petals; sepals 5, united below; petals 5, united
below to form a short tube; stamens 5, attached to the corolla
tube alternate with its lobes; pistil 1; style terminal, 2-cleft;
ovary 1-celled or nearly 2-celled; fruit a capsule, opening by
2 valves, seeds few to several.
Hydrophyllum fendleri (Gray) Heller - Fendler Waterleaf
Phacelia denticulata Osterh. - Phacelia
Phacelia hastata Dougl. ex Lehm. - Phacelia
31. IRIDACEAE Iris Family (page 184)
Herbs; leaves 2-ranked; flowers perfect, regular, blue or purple;
sepals 3, petaloid; petals 3; stamens 3; pistil 1, of 3 united
carpels; style branches sometimes appearing like 3 additional
petals; ovary inferior, 3-celled; fruit a loculicidal capsule.
Sisyrinchium angustifolium Mill. - Common Blue-eyed Grass
Sisyrinchium montanum Greene - Montane Blue-eyed Grass
32. JUNCACEAE Rush Family (page 185)
Herbs; leaves simple, linear or filiform, basal or alternate,
sheathing; flowers bisexual, regular, in a panicle, cyme, or head,
rarely solitary; perianth of 6, mostly similar, scale-like, separate
parts; stamens 3 or 6; pistil 1; ovary superior; styles 1 or 3;
locules 1 or 3; fruit a capsule.
Juncus balticus Willd. var. montanus Engelm. - Baltic Rush
Juncus bufonius - Toad Rush
Juncus ensifolius - Sword-Leaf Rush
Juncus longistylis - Long-Styled Rush
Juncus tenuis - Slender Rush
33. LAMIACEAE Mint Family (page 188)
Aromatic herbs with a minty odor; stems square in cross section;
leaves simple and opposite; flowers hypogynous, with united petals;
corolla nearly regular to irregular, often 2-lipped; stamens 4,
in two sets of 2 each, or only 2, attached to the corolla tube
alternate with its lobes; pistil 1; ovary deeply 4-lobed, style
1, arising from between the lobes of the ovary, 2-forked at the
summit; fruit a group of four 1-seeded nutlets.
Agastache urticifolia - Nettle-Leaved Giant Hyssop
Dracocephalum parviflorum Nutt. - American Dragonhead
Mentha arvensis - Common Mint
Monarda fistulosa L. var. methaefolia (Grah.) Fern. - Wildbergamot, Beebalm
Scutellaria brittonii Porter - Brittons Skullcap
Scutellaria galericulata L.var. brittonii Porter - Skullcap
Stachys palustris - Water Hedge Nettle
34. LILIACEAE Lily Family (page 191)
Herbs or sometimes shrubs; flowers mostly showy; perianth consisting
of 3 sepals and 3 petals that are usually similar and all petaloid;
sepals not green; stamens 6; pistil 1, of 3 united carpels; styles
separate or united; ovary superior, 3-celled and with several
to many ovules; fruit a capsule or sometimes a berry.
Allium cernuum Roth. - Nodding Onion
Allium textile Nels. &Macbr. - Prairie Onion
Calochortus gunnisonii Wats. - Gunnison Mariposa Lily
Leucocrinum montanum Nutt. ex Gray - Common Star Lily
Smilacina stellata (L.) Desf. - Starry False Solomon's Seal
Yucca glauca - Spanish Bayonet
35. LOASACEAE Loasa Family (page 196)
Annual or perennial herbs with barbed hairs; leaves simple, entire
to pinnatifid, alternate; flowers bisexual, regular, in cymes
or rarely solitary; calyx lobes mostly 5, borne on a short hypanthium;
petals mostly 5, yellow to whitish, separate, sometimes also with
an inner whorl of 3 or more (mostly 5) petal-like staminodia;
stamens many; pistil 1; ovary inferior; style 1; locule 1; ovules
many; fruit a capsule.
Mentzelia dispersa Wats. - Bushy Mentzelia
Mentzelia rusbyi Wooton - Desert Mentzelia
36. NYCTAGINACEAE Four-O'Clock Family (page 200)
Herbs; leaves usually opposite, simple, mostly entire; flowers
bisexual or rarely unisexual, regular, each flower or a cluster
of flowers subtended by usually 5 separate or united bracts (involucre),
the involucres in a cyme or panicle; calyx of 5 united petaloid
sepals; corolla none; stamens mostly 3-5; pistil 1; ovary superior
but often appearing superficially inferior by constriction of
the calyx tube; style 1; fruit an achene, sometimes enclosed by
the calyx.
Mirabilis hirsuta Pursh - Hairy Four-O-Clock
Mirabilis nyctaginea - Heart-Leaved Umbrellawort
37. ONAGRACEAE Evening Primrose Family (page 201)
Herbs; flowers regular, epigynous, often with a conspicuous, tubular
hypanthium extending above the ovary, flowers often large and
showy, variously colored but often white or pink; sepals 4; petals
4; stamens 4 or 8, attached to the rim of the hypanthium or to
the top of the ovary when the hypanthium is lacking; pistil 1;
style single; stigma often 4-lobed; ovary inferior, usually 4-celled;
fruit a capsule or sometimes nutlike and indehiscent.
Epilobium anagallidifolium - Dwarf Willow Herb
Epilobium angustifolium - Fireweed
Epilobium ciliatum Raf. - Cilliate Willow Herb
Epilobium watsonii Raf. - Purpleleaf Willowherb
Gaura coccinea Nutt. ex Pursh - Scarlet Gaura
Gayophytum diffusum T.&G.var. diffusum - Bigflower Groundsmoke
Oenothera caespitosa Nutt. - Tufted Evening Primrose
Oenothera coronopifolia T &G - Evening Primrose
Oenothera depressa Greene - Evening Primrose
38. ORCHIDACEAE Orchid Family (page 207)
Herbs; flowers perfect, irregular, epigynous; sepals 3, similar
and petaloid; petals 3, dissimilar, one of them projecting downward
and of a different shape (the lip); stamens 1 or 2, attached near
the end of the style; anther or anthers sessile; pistil of 3 united
carpels; ovary inferior and 1-celled; parietal placentae; fruit
a 1-celled capsule.
Corallorhiza maculata - Maculate Coral Root
Habenaria viridis - Green Bog Orchid
39. OROBANCHACEAE Broomrape Family (page 209)
Herbs without green color, brownish or yellowish; parasitic on
the roots of other plants; leaves scale-like; flowers hypogynous,
irregular; sepals 5, united; petals 5, united; stamens 4, in two
sets of 2 each, attached to the corolla tube alternate with its
lobes; pistil 1; ovary 1-celled; parietal placentation; fruit
a many-seeded capsule.
Orobanche fasciculata Nutt. - Purple Broomrape
40. PINACEAE Pine Family (page 28)
Resinous, evergreen trees or shrubs; leaves needle-like; cones
woody; seeds produced in pairs at the base of the cone scales
(sporophylls), not enclosed in an ovary or pod.
Picea engelmannii - Engelmann Spruce
Pinus flexilis James - Limber Pine
Pinus ponderosa Laws. &Laws. - Ponderosa Pine
41. PLATANGINACEAE Plantain Family (page 210)
Herbs; leaves in basal clusters, simple, entire, parallel-veined;
flowering stems (scapes) naked and ending in a dense spike of
small flowers; flowers hypogynous, regular; sepals 4, united;
petals 4, spreading, scarious and whitish, united below into a
short tube; stamens 2 or 4, attached to the corolla tube alternate
with its lobes; pistil 1; ovary 1-4-celled; fruit usually circumscissile
capsule (pyxis).
Plantago patagonica Jacq. var. spinulosa (Decne) Gray - Woolly Plantain
42. POACEAE Grass Family (page 211)
Herbs; leaves simple, alternate, sheathing, parallel-veined, 2
ranked, with a hairy or membranous ligule at junction of sheath
and blade on inner side; flowers bisexual or unisexual, arranged
in spikelets, each spikelet usually consisting of 2 empty lower
bracts (glumes) subtending 1 or more florets, each floret composed
of 2 bracts (a lemma which usually has a midnerve, and a palea
which usually lacks a midnerve) which subtend the flower; perianth
greatly reduced; stamens 1-6, usually 3; ovary 1, superior; styles
1-3, usually 2; fruit a caryopsis.
Agropyron caninum - Slender Wheatgrass
Agropyron cristatum - Crested Wheatgrass
Agropyron dasystachyum (Hook.) Scribn. - Thickspine Wheatgrass
Agropyron elongatum - Tall Wheatgrass
Agropyron smithii Rydb. - Western Wheatgrass
Agropyron spicatum - Bluebunch Wheatgrass
Alopercurus aequalis Sobel. - Water Foxtail
Bouteloua gracilis (H.B.K.) Lag. ex Griffiths - Blue Grama
Bromus ciliatus - Ciliate Bromegrass
Bromus tectorum L. - Cheatgrass Brome
Catabrosa aquatica - Brookgrass
Cinna latifolia - Woodreed
Danthonia parryi Scribn. - California Danthonia
Festuca idahoensis Elmer - Idaho Fescue
Glyceria grandis - Big Mannagrass
Glyceria striata - Striate Mannagrass
Hordeum jubatum L. - Foxtail Barley
Koeleria macrantha (Ledeb.) Schult. - Prairie Junegrass
Leucopoa kingii - Western Grass
Muhlenbergia filiculmis Vasey - Slimstem Muly
Muhlenbergia filiformis - Pullup Muhly
Oryzopsis micrantha (Trin. &Rupr.) Thurb. - Littleseed Ricegrass
Phleum pratense L. - Common Timothy
Poa compressa L. - Canada Bluegrass
Poa cusickii Vasey var. cusickii - Cusick Bluegrass
Poa fendleriana (Steud) var. nervosa (Hook.) Vasey - Mutton Bluegrass
Poa juncifolia - Big Bluegrass
Poa nervosa - Nerved Bluegrass
Poa pratensis L. - Kentucky Bluegrass
Poa sandbergii Vasey - Sandberg Bluegrass
Poa scabrella Thurb. - Pine Bluegrass
Puccinellia nuttalliana - Nuttall Alkali Grass
Sitanion hystrix Nutt. - Bottlebrush Squirreltail
Stipa columbiana - Columbian Needlegrass
Stipa comata Trin &Rupr. var. comata - Needle and Thread
Stipa viridula - Green Needlegrass
43. POLEMONIACEAE Phlox Family (page 234)
Low to moderately tall herbs, rarely shrubby; leaves alternate
or opposite; sepals 5, united into a tube below; petals 5, united
into a tube below; stamens 5, attached to the slender lobes; ovary
3 celled, with axillary placentation; fruit a 3-valved capsule.
Collomia linearis Nutt. - Narrowleaved Collomia
Microsteris gracilis (Hook.) Greene var. humilior (Hook.) Cronq. - Microsteris
44. POLYGONACEAE Buckwheat Family (page 238)
Herbs, often with sour juice; leaves alternate and entire; stems
often papery stipular sheaths at the joints (except inEriogonum which has no stipules); flowers small, solitary in the axils
of leaves, in terminal, compact racemes, in umbels, in capitate
clusters, or paniculate, perfect or sometimes unisexual; perianth
of 5-6 parts, sometimes united, petaloid and in 1 series or sepaliod
and in 2 series, often persistent and sometimes becoming greatly
enlarged in fruit; stamens 4-12; pistil 1; ovary superior; styles
2 or 3; stigmas 2 or 3; fruit a lenticular or triangular achene.
Eriogonum alatum Torr. - Wing Wild Buckwheat
Eriogonum jamesii Benth. - James Wildbuckwheat
Eriogonum umbellatum (Torr.) var. majus Hook. - Sulfur Wild Buckwheat
Polygonum aviculare - Creeping Knotweed
Polygonum convolvulus - Twining Knotweed
Polygonum douglasii Greene var. microspermum Englein. - Douglas Knotweed
Rumex crispus - Crispy Dock
Rumex salicifolius - Willow-Leaved Dock
Rumex utahensis Rech - Dock
45. POLYPODIACEAE Fern Family (page 24)
Herbs; leaves well developed, often lobed or dissected; reproduction
by spores in minute spore case (sporangia) that are clustered
in dots, lines or variously shaped clusters (sori) on the backs
or margins of the leaves, or sometimes on specialized leaves.
Cystopteris fragilis (L.) Bernh. - Brittle Bladderfern
Woodsi oregana - Brittle Fern
46. PORTULACACEAE Purslane Family (page 244)
Succulent herbs; leaves alternate, opposite, or basal, simple
and entire; flowers solitary or racemose, regular, perfect, small
or showy; sepals often 2, sometimes more, persistent and often
enlarged in fruit; petals usually 5, sometimes more, white to
pink to rose; stamens often 5, sometimes more; pistil 1, of 2
or more united carpels; ovary usually superior, 1-celled, free-central
placentation; fruit a capsule with numerous seeds.
Claytonia lanceolata Pursh var. flava (A. Nels.) Hiteho. - Lanceleaf Springbeauty
Lewisia pygmaea (Gray) Robins - Least Lewisia
Montia chamissoi (Ledeb. ex Spreng.) Greene - Chamissa Minerslettuce
47. PRIMULACEAE Primrose Family (page 247)
Herbs; leaves simple, basal or sometimes cauline; flowers hypogynous,
regular, with united petals; sepals 5, partly united; petals 5,
united into a tube below; stamens 5, attached to the corolla opposite
its lobes; pistil 1, with a single and stigma; ovary superior,
1-celled, free-central placentation; fruit a capsule.
Androsace filiformis - Filiform Rockjasmine
Androsace septentrionalis L. - Pygmyflower, Rock Jasmine
Dodecatheon pulchellum (Raf.) Merr. - Shootingstar
48. RANUNCULACEAE Buttercup Family (page 249)
Herbs; flowers hypogynous, the parts all separate from one another;
perianth regular or irregular; sepals 3 or more, often petaloid
when the petals are lacking; petals none or several; stamens numerous,
spirally arranged; pistils (carpels) 3 or more, each with 1 or
several ovules; fruit a group of achenes or follicles, or sometimes
a berry.
Aconitum columianum - Columbian Monkshood
Actaea rubra - Red Baneberry
Anemone canadensis L. - Canada Anemone
Anemone multifida Poir - Cutleaf Anemone
Anoemone patens L. - Spreading Pasqueflower
Clematis ligusticifolia Nutt. - Westernwhite Clematis
Delphinium geyeri - Geyer Larkspur
Delphinium nuttallianum Pritz - Larkspur
Delphinium occidentale - Mountain Larkspur
Ranunculus abortivus - Premature Buttercup
Ranunculus acriformus - Sharp Buttercup
Ranunculus cymbalaria - Shore Buttercup
Ranunculus glaberrimus Hook. var. ellipticus Greene - Sagebrush Buttercup
Ranunculus gmelinii - Gmelin Buttercup
Ranunculus macounii - Macoun Buttercup
Ranunculus ranunculinus (Nutt.) Rydb. - Nuttail Buttercup
Ranunculus sceleratus - Hollow-Stem Buttercup
Thalictrum fendleri Engelm. - Fendler Meadowrue
Thalictrum occidentale - Western Meadow Rue
Thalictrum sparsiflorum - Scattered-Flower Meadow Rue
Thalictrum venulosum - Veiny Meadow Rue
49. ROSACEAE Rose Family (page 259)
Herbs, shrubs, or trees; leaves alternate, simple or compound,
often with prominent stipules; flowers regular, perigynous, sometimes
appearing to be epigynous; sepals, petals, and stamens inserted
on the rim of a shallow or deep hypanthium that is free from the
pistil or pistils or that encloses the pistil or pistils and is
sometimes adnate to the pistil; sepals 5; petals usually 5, sometimes
none; stamens usually numerous; pistil 1 and of 1 or several united
carpels, or the pistils and carpels several to many and separate;
fruit a group of achenes, a group of follicles, an aggregate of
achenes (strawberry), an aggregate of druplets (raspberry), a
single drupe (cherry), a single pome (apple), or an urn-shaped
structure enclosing several achenes (rose hip).
Amelanchier alinfolia Nutt. - Saskatoon Serviceberry
Cercocarpus montanus Raf. - True Mountain Mohogany
Fragaria virginiana Miller - Virginia Strawberry
Geum macrophylum - Big-Leaf Avens
Physocarpus monogynus (Torr.) Coult. - Mountain Ninebark
Potentilla concinna Richards var. concinna - Elegant Cinquefoil
Potentilla fissa Nutt. - Bigflower Cinquefoil
Potentilla gracilis Dougl. ex Hook. var. nuttallii Lehm. - Showy Cinquefoil
Potentilla hippiana Lehm. - Horse Cinquefoil
Potentilla norvegica L. - Norwegian Cinquefoil
Potentilla pensylvanica L. - Pennsylvania Cinquefoil
Prunus virginiana L. var. melanocarpa (A. Nels.) Sarg. - Common Chokecherry
Purshia tridentata (Pursh) DC - Antelope Bitterbrush
Rosa sayi Schwein - Rose
Rosa woodsii Lindl. - Woods Rose
Rubus idaeus L. - Red Raspberry
50. RUBIACEAE Madder Family (page 267)
Herbs; stems square; leaves entire, simple, whorled; stems often
with fine, retrorse prickles; flowers epigynous, regular, small,
white; calyx with 4 small to minute lobes; corolla of united petals,
4 lobed, the lobes spreading and the tube short; stamens usually
4, attached to the tube of the corolla alternate with its lobes;
pistil 1, of 2 carpels, each carpel rounded and sometimes with
barbed bristles; fruit a pair of 1-seeded nutlets.
Galium boreale L. - Northern Bedstraw
Galium trifidum - Trifid Bedstraw
Galium triflorum - Three-Flower Bedstraw
51. SALICACEAE Willow Family (page 268)
Tree or shrubs; leaves alternate, simple, stipulate; leaf margins
serrate, denate, or sometimes lobed; inflorescence a catkin, flowers
unisexual, dioecious, lacking perianth; seeds numerous and each
with a tuft of cottony hairs.
Populus angustifolia - Narrowleaf Cottonwood
Populus tremuloides Michx. - Quaking Aspen
Salix bebiana - Bebb Willow
Salix exigua - Coyote Willow
Salix lasiandra - Whiplash Willow
Salix lemmonii - Lemmon Willow
Salix monticola - Mountain Willow
Salix planifolia - Plane-Leaved Willow
Salix scouleriana - Scouler Willow
52. SANTALACEAE Sandalwood Family (page 275)
Partially parasitic, perennial herbs; pale green color, with root
connections to other plants; stems arising from horizontal rhizomes;
leaves alternate, entire, glaucous; flowers small and greenish
white; inflorescences umbel-like cymes; sepals 5; petals none;
stamens 5, opposite the sepals and attached to a disk at the summit
of the ovary; pistil 1; ovary inferior; fruit a purple, 1-seeded,
drupe like berry crowned by the persistent calyx.
Comandra umbellata (L.) Nutt. var. pallida (A. DC) Jones - Common Bastard Toadflax
53. SAXIFRAGACEAE Saxifrage Family (page 275)
Herbs; leaves alternate or basal without stipules; leaf blades
often but not always with palmate venation; flowers perigynous;
sepals, petals, and the stamens inserted on the rim of a hypanthium;
sepals usually 5; petals usually 5, stamens usually 5 or 10; carpels
2, more or less united in a single pistil that is 2-horned at
the apex; ovary either 1-celled or 2-celled; fruit a capsule.
Heuchera bracteata (Torr.) Ser. - Alumroot
Heuchera parvifolia Nutt. ex T.&G. - Littleleaf Alumroot
Lithophragma glabrum Nutt. var. ramulosum (Suksd.) Boivin - Woodland Star
Saxifraga rhomboidea Greene - Diamondleaf Saxifrage
54. SCROPHULARIACEAE Figwort Family (page 278)
Herbs; leaves alternate or opposite; stems nearly always rounded
in cross section; flowers hypogynous; petals united; corolla irregular,
often 2-lipped; stamens 4, in two sets of 2 each, often with a
fifth sterile filament, all attached to the corolla tube alternate
with its lobes; pistil 1; ovary not lobed; style single, not bifid
at the apex; fruit a 2-celled capsule.
Besseya wyomingensis (A. Nels.) Rydb. - Wyoming Kittentails
Castilleja flava - Yellow Indian Paintbrush
Castilleja linariaefolia Benth. - Indian Paintbrush
Collinsia parviflora Lindl. - Blue-eyed Mary
Linaria dalmatica - Dalmation Toadflax
Orthocarpus luteus Nutt. - Yellow Owlclover
Pedicularis procera - Tall Lousewort
Penstemon glaber Pursh var. alpinus (Torr.) Gray - Alpine Penstemon
Penstemon secundiflorus Benth. - Sidebells Penstemon
Penstemon virens Penn. ex Rydb. - Green Penstemon
Scrophularia lanceolata Pursh - Figwort
Verbascum thapsus L. - Flannel Mullein
Veronica americana Schwein ex Benth. - American Speedwell
Veronica peregrina - Wandering Speedwell
55. SELAGINELLACEAE Clubmoss Family (page 27)
Low, mosslike herbs; stems branching and unjointed, covered with
small, awl-shaped, spirally arranged leaves; reproduction by spores
of two sizes, produced on sporophylls that form terminal cones;
found mainly in rocky, hilly places.
Selaginella densa - Rock Spike Moss
56. SOLONACEAE Nightshade Family (page 288)
Annual herbs in disturbed areas; leaves mostly alternate, simple,
or rarely compound; flowers bisexual, regular or slightly irregular,
solitary; calyx 5 lobed; corolla blue, violet, white, or yellow;
stamens 5, attached to the corolla and alternate with the lobes;
pistil 1, ovary superior; style 1; locules 2; ovules many; fruit
a berry.
Solanum triflorum Nutt. - Cutleaf Nightshade
57. SPARGINACEAE Bur-Reed Family (page 289)
Monoecious aquatic herbs; leaves simple, linear, alternate; staminate
flowers in globose heads above globose heads of pistillate flowers;
perianth reduced to usually 3-6 membranous scales; stamens 3 or
more; ovary superior; styles 1 or 2; fruit a nutlet or achene.
Sparganium emersum - Emersed Bur-Reed
58. URTICACEAE Nettle Family (page 291)
Pubescent herbs; leaves simple, alternate or opposite, petioled;
flowers unisexual or imperfectly bisexual, mostly regular, in
a cyme, panicle, or spike, the plants monoecious or dioecious;
perianth greenish or brownish, of 4 separate or united parts,
sometimes 2 smaller than the other 2; stamens mostly 4; pistil
1; ovary superior; style 1; fruit an achene.
Parietaria pensylvanica Muhl. ex Willd. - Pennsylvania Pellitory
Urtica dioica L. - Stinging Nettle
59. VALERIANACEAE Valerain Family (page 291)
Herbs with malodorous roots; leaves opposite; flowers small and
often numerous, white, epigynous, regular or somewhat irregular,
the corolla of united petals; sepals 5; petals 5, united into
a tube; stamens 1-3, attached to the tube of the corolla; pistil
1; ovary inferior; fruit 1-celled and 1-seeded, indehiscent, with
a late-developing calyx of several feathery bristles (pappus)
at the summit.
Valeriana edulis - Edible Valerian
60. VERBANACEAE Verbena Family (page 292)
Annual to perennial herbs; leaves simple or compound, usually
opposite; flowers mostly bisexual, irregular or rarely regular,
usually in head-like or elongate bracteate spikes; calyx usually
2-5 lobed; corolla usually unequally 4 or 5 lobed; stamens usually
4; pistil 1; ovary superior, often 2 or 4 lobed; style 1; locules
2 or 4; ovules usually 1 per locule; fruit 2 or 4 nutlets.
Verbena bracteata - Creeping Verbena
61. VIOLACEAE Violet Family (page 292)
Low herbs; leaves alternate or basal, stipulate, blades narrowly
lanceolate to oblong or cordate; flowers hypogynous or somewhat
perigynous, the corolla irregular, the lower petal saccate or
spurred; sepals 5; petals 5; stamens 5, the anthers meeting but
not united over the pistil; ovary 1 celled, with 3 parietal placentae;
fruit a 3-valved capsule.
Viola canadensis L. - Canada White Violet
Viola nephrophylla - Stemless Blue Violet
LITERATURE CITED
?Dorn, R. D. 1988. Vascular plants of Wyoming. Mountain West Publ.:
Cheyenne, Wyoming.
?Porter, C. L. 1968. Spring flora of southeastern Wyoming. University
of Wyoming Agricultural Experiment Station, Bulletin #491.